Jonathan, the world's oldest known tortoise, has passed away at the age of 193, leaving behind a legacy of resilience and longevity. The giant tortoise had been cared for by a veterinarian who described his passing as a loss for the animal kingdom. Jonathan's remarkable life has inspired many, and his memory will live on through his remarkable story.

The vet who cared for the giant tortoise said he had left behind a "legacy of resilience and longevity".